Ibaraki University's
Graduate School of Science and Engineering (Science)
Department of Science (Biological Sciences Course)



  • Birth:1969

Academic background

  1. The University of Tokyo Faculty of Arts and Sciences 基礎科学科第二 1991/03 Graduated
  2. The University of Tokyo Graduate School of Arts and Sciences 広域科学 Doctor course 1996/03 Completed

Academic degrees

  1. 博士(学術) The University of Tokyo 1996/03

Research Areas

  1. Molecular Phylogenetics
  2. Ecology

Research keywords

  1. 動物生態学・系統学

Subject of research

  1. Ecological and phylogenetic study on termites and symbiotic protists シロアリと共生原生生物群集の進化過程を系統推定手法を用いて明らかにする
  2. シロアリのカスト決定機構に関する生態・進化学的研究 シロアリのカスト決定における遺伝的基盤と、その進化過程・生態学的意義を明らかにすることを目指す


  1. 行動生物学事典 東京化学同人 2013/11/22 978-4-8079-0837-0
  2. シロアリの事典 海青社 2012/12/01 978-4-86099-260-6
  3. 生態学事典 巌佐庸・松本忠夫・菊沢喜八郎(編) 共立出版 2003/06/25 4-320-05602-7
  4. Termites: Evolution, Sociality, Symbioses, Ecology. T. Abe et al. (eds.) Kluwer Academic Publishers 2000
  5. シロアリと微生物 「住まいとシロアリ」 海青社 2000


  1. Research paper (scientific journal) Joint Host-symbiont cospeciation of termite-gut cellulolytic protists of the genera Teranympha and Eucomonympha and their Treponema endosymbionts. Noda S, Shimizu D, Yuki M, Kitade O, Ohkuma M Microbes Environ. 33/ 1, 26-33 2018 10.1264/jsme2.ME17096
  2. Research paper (scientific journal) Joint シロアリの腸内共生原生生物群集の特性と機能 北出 理, 野田 悟子 生物科学 68/ 3, 154-164 2017
  3. Research paper (scientific journal) Joint Intracolonial genetic variation affects reproductive skew and colony productivity during colony foundation in a parthenogenetic termite Miyazaki S, Yoshimura M, Saiki R, Hayashi Y, Kitade O, Tsuji K, Maekawa K BMC Evolutionary Biology 14, 177 2014 10.1186/s12862-014-0177-0
  4. Research paper (scientific journal) Joint Neotenic reproductives influence worker caste differentiation in the termite Reticulitermes speratus (Isoptera; Rhinotermitidae). Hayashi, Y., Miyata, H., Kitade, O., Lo, N. Sociobiology 60/ 4, 446-452 2013
  5. Research paper (scientific journal) Joint Symbiotic protosit communities in the termite Coptotermes formosanus in Japan and a comparison of community structures between workers and soldiers. Kitade O., Hayashi, Y., and Noda, S. Japanese Journal of Protozoology 46, 21-29 2013


  1. 日本昆虫学会賞 1999

Memberships of academic societies

  1. 日本昆虫学会
  2. 国際社会性昆虫学会(International Union for Study of Social Insects)
  3. 日本原生生物学会 1995-Present
  4. 日本生態学会 1992-Present